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[\'Zeeman Institute for Systems Biology and Infectious Disease Epidemiology Research, Mathematics Institute, and School of Life Sciences, University of Warwick, Coventry, UK.\', \'Zeeman Institute for Systems Biology and Infectious Disease Epidemiology Research, Mathematics Institute, and School of Life Sciences, University of Warwick, Coventry, UK. Electronic address: m.j.keeling@warwick.ac.uk.\']
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